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Copepoda (excl. Harpacticoida)
Boxshall, G. (2001). Copepoda (excl. Harpacticoida), in: Costello, M.J. et al. European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 252-268
In: Costello, M.J.; Emblow, C.; White, R.J. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50. Muséum national d'Histoire naturelle: Paris. ISBN 2-85653-538-0. 463 pp., more
In: Collection Patrimoines Naturels. Muséum national d'Histoire naturelle: Paris. ISSN 1158-422X, more

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    Copepoda [WoRMS]

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  • Boxshall, G., more

    The ordinal level classification of the copepods is based on Huys & Boxshall (1991). Data on the pelagic copepods were drawn primarily from the two excellent volumes on the diversity and distribution of pelagic copepods published by Claude Razouls (1995, 1996). These data were supplemented with the new taxa recorded or described from European waters since 1996 (Fosshagen et al., in press; Jaume & Boxshall, 1995, 1996a-d, 1997a-b; Jaume et al., 1999; Jaume et al., in press; Markhaseva, 1996; Zagami et al., 2000) and updated to include recent changes in nomenclature (Belmonte, 1998; Böttger-Schnack, 1999; Boxshall & Jaume, in press; Huys & Böttger-Schnack, 1997). Included in these papers are the many new taxa that have been discovered in anchialine cave habitats in the Mediterranean region and in the Canary Islands.

    The parasitic copepods of marine fishes have been compiled primarily from the following major sources: the monograph of Kabata (1979 and the analysis of species richness of parasitic copepods of Mediterranean fishes by Raibaut, Combes & Benoit (1998). Other changes and new species are included (Boxshall, in press; Hogans, 1987). The copepods associated with marine invertebrate hosts in Northern European waters were summarised by Gotto (1993). Data for copepods from invertebrates in Southern European waters and the Mediterranean region were widely scattered through the primary literature and are probably only 90 to 95% complete. Data has been drawn from standard references (e.g. Giesbrecht, 1899; Illg & Dudley, 1961, 1965, 1980) and supplemented by information from subsequent publications (Boxshall & Defaye, 1995; Ho et al., 1998; Holmes, 1998; Humes & Boxshall, 1996; Lamb et al., 1996; López-González, et al., 1998; López-González & Conradi, 1996; Marchenkov & Boxshall, 1995; O’Reilly, 1995). The order Monstrilloida has been the subject of profound nomenclatural change (see Grygier, 1995 and 1997).Additional data and corrections were provided by Myles O’Reilly and Dusan Zavodnik.


    • Apostolov, A. & T.M. Marinov, 1988. Copepoda Harpacticoida (morski kharpaktikoidi). [Copepoda, Harpacticoida (marine harpacticoids)]. - Fauna Bolgarii [Fauna Bulgarica], 18: 1-384. Sofia, Izd. B'lg. Akad. Nauk. (Bulgarian).
    • Belmonte, G. 1998. Pteriacartia, a new genus of Acartiidae (Calanoida, Diaptomoidea) for Acartia josephinae Crisafi, 1974. J. Mar. Syst. 15: 359-368.
    • Bodin, P., 1964. Recherches sur la systématique et la distribution des Copépodes Harpacticoïdes des substrats meubles des environs de Marseille. - Recl Trav. Stn mar. Endoume, 51 (=Bull. 35): 107-183.
    • Böttger-Schnack, R. 1999. Taxonomy of Oncaeidae (Copepoda, Poecilostomatoida) from the Red Sea. – I. 11 species of Triconia gen nov. and a redescription of T. similis (Sars) from Norwegian waters. Mitt. Hamb. Zool. Mus. Inst. 96: 37-128.
    • Boxshall, G.A. (in press) Parasitic copepods (Copepoda: Siphonostomatoida) from deep-sea and mid-water fishes. Systematic Parasitology
    • Boxshall, G.A. & Jaume, D. (in press) Discoveries of Cave Misophrioids (Crustacea: Copepoda) Shed New Light on the Origin of Anchialine Faunas. Zool. Anz.
    • Boxshall, G.A. & Defaye, D. 1995 Copépodes nouveaux (Siphonostomatoida: Nicothoidea) parasites de Cumacés et des Décapodes profonds. Bull. natn. Mus. Hist. nat. 17: 283-296.
    • Fosshagen, A., Boxshall, G.A. & Iliffe, T.M. (in press) The Epacteriscidae, a cave-living family of calanoid copepods. Sarsia
    • Giesbrecht, W. 1899. Die Asterocheriden des Golfes von Neapel und der angrezenden Meeres-Abschnitte. Fauna Flora Golf. Neapel 25: i-vi, 1-217, pls 1-11.
    • Gotto, V. 1993. Commensal and Parsitic Copepods associated with marine invertebrates (and whales). In: D.M. Kermack, R.S.K. Barnes & J.H. Crothers eds,
    • Synopses of the British Fauna (New Series), No. 46: 1-264.Grygier, M.J. 1995. Annotated chronological bibliography of Monstrilloida (Crustacea: Copepoda). Galaxea 12: 1-82.
    • Grygier, M.J. 1997. Monstrilla Dana, 1849 and Thaumaleus Krøyer, 1849 (Crustacea, Copepoda): conserved. Bull. zool. Nom. (Opinion 1869) 54 (2): 131-132.
    • Ho, J.-S., Conradi, M. & López-González, P.J. 1998. A new family of cyclopoid copepods (Fratiidae) parasitic in the ascidian (Clavelina dellavallei) from Cádiz, Spain. J. Zool., Lond. 246: 39-48
    • Hogans, W.E. 1987. Morphological variation in Pennella balaenoptera and P. filosa (Copepoda: Pennellidae) with a review of the genus Pennella Oken, 1816 parasitic on Cetacea. Bull. mar. Sci. 40 (3): 442-453.
    • Holmes, J.M.C. 1998. Phenetic relationships among some free-living siphonostomatoid copepods from southwest Ireland, and the description of Glannapontius maculatus gen. et sp. nov. J. Marine Systems 15: 229-241.
    • Humes, A.G. & Boxshall, G.A. 1996 A revision of the lichomolgoid complex (Copepoda: Poecilostomatoida), with the recognition of six new families. J. nat. Hist. 30: 175-227.
    • Huys, R., 1996. Superornatiremidae fam. nov. (Copepoda: Harpacticoida): An enigmatic family from North Atlantic anchihaline caves. - Sci. Mar., 60(4): 497-542.
    • Huys, R. & Böttger-Schnack, R. 1997. On the diphyletic origin of the Oncaeidae Giesbrecht, 1892 (Copepoda: Poecilostomatoida) with a phylogenetic analysis of the Lubbockiidae fam. nov. Zool. Anz. 235 (3-4): 243-261.
    • Huys, R. & Boxshall, G.A. 1991. Copepod Evolution: 1-468. The Ray Society, London.Huys, R., J.M. Gee, C.G. Moore & R. Hamond, 1996. Marine and brackish water harpacticoid copepods. Part 1.
    • Huys, R. & T.M. Iliffe, 1998. Novocriniidae, a new family of harpacticoid copepods from anchihaline caves in Belize. Zool. Scr., 27: 1-15.
    • Illg, P.L. & Dudley, P.L. 1961. Notodelphyid copepods from Banyuls-sur-Mer. Vie Milieu (Supplement 12): 1-126.
    • Illg, P.L. & Dudley, P.L. 1965. Notodelphyid copepods from the vicinity of Naples. Pubbl. Staz. zool. Napoli 34: 373-451.
    • Illg, P.L. & Dudley, P.L. 1980. The family Ascidicolidae and its subfamilies (Copepoda, Cyclopoida), with descriptions of new species. Mém. Mus. Hist. nat. Paris (N.S.) (A) (Zoologie) 117: 1-192.
    • Jaume, D. & Boxshall, G.A. 1995 Stygocyclopia balearica, a new genus and species of calanoid copepod (Pseudocyclopiidae) from anchihaline caves in the Balearic Islands (Mediterranean). Sarsia 80: 213-222.
    • Jaume, D. & Boxshall, G.A. 1996a A new genus and two new species of cave-dwelling copepods from the Balearic Islands (Mediterranean). J. nat. Hist. 30: 989-1006.Jaume, D. & Boxshall, G.A. 1996b Rare cyclopoid copepods (Crustacea) from Mediterranean littoral caves. Bull.nat.Hist.Mus.Lond. (Zool.) 62: 83-99.
    • Jaume, D. & Boxshall, G.A. 1996c Two new genera of cyclopinid copepods (Crustacea) from anchihaline caves on western Mediterranean Islands. Zool. J. Linn. Soc. 117: 283-304.
    • Jaume, D. & Boxshall, G.A. 1996d The persistence of an ancient marine fauna in Mediterranean waters: new evidence from misophrioid copepods living in anchihaline caves. J. nat. Hist. 30: 1583-1595.
    • Jaume, D. & Boxshall, G.A. 1997a A new genus and two new species of misophrioid copepods (Crustacea) from the North Atlantic hyperbenthos. Sarsia 82: 39-54.
    • Jaume, D. & Boxshall, G.A. 1997b Two new genera of cyclopinid copepods (Cyclopoida: Cyclopinidae) from anchihaline caves of the Canary and Balearic islands, with a key to genera of the family. Zool. J. Linn. Soc. 120: 79-101.
    • Jaume, D, Cartes, J.E. & Boxshall, G.A. (in press) Shallow-water and not deep-sea as most plausible origin for cave-dwelling Paramisophria species (Copepoda: Calanoida: Arietellidae), with description of three new species from Mediterranean bathyal hyperbenthos and littoral caves. Contrib. Zool.
    • Jaume, D. Fosshagen, A. & Iliffe, T.M. 1999. New cave-dwelling pseudocyclopiids (Copepoda, Calanoida, Pseudocyclopiidae) from the Balearics, Canary, and Philippine archipelagos. Sarsia 84: 391-417.
    • Kabata, Z. 1979. Parasitic Copepoda of British fishes. No. 152: i-xii, 1- 468, figs 1-2031. The Ray Society, London.
    • Lamb, E.J., Boxshall, G.A., Mill, P.J. & Grahame, J. 1996 A new family of endoparasitic copepods (Poecilostomatoida) from the Dog Whelk Nucella lapillus. J. Crustacean Biol. 16: 142-148.
    • Lang, K., 1948. Monographie der Harpacticiden, I: 1-896, figs. 1-361; II: 897-1682, figs. 362-607, maps 1-378.
    • López-González, P.J., Bresciani, J. & Conradi, M. 1998. Tarificola bulbosus, new genus, new species, a highly transformed parasitic copepod, with information on its parasitism and larval development. J. crust. Biol. 18 (3): 581-589.
    • López-González, P.J. & Conradi, M. 1996. Mychophilus palmatus, a new species (Copepoda: Cyclopoida: Ascidicolidae) associated with the compound ascidian Botryllus leachi Savigny, with remarks on congeneric species and related genera. Hydrobiologia 330: 67-72.
    • Marchenkov, A.V. & Boxshall, G.A. 1995 A new family of copepods associated with ascidiaceans in the White Sea, and an analysis of antennulary segmentation and setation patterns in the order Poecilostomatoida. Zool. Anz. 234: 133-144.
    • Markhaseva, E.L. 1996. Calanoid copepods of the family Aetideidae of the World Ocean: 1-331. St Petersburg. ( Trudy Zoologicheskogo Instituta RAN 268).
    • Noodt, W., 1955. Marmara denizi Harpacticoid'Leri (Crust. Cop.). Marine Harpacticoiden (Crust. Cop.) aus dem Marmara Meer. - Istanb. Univ. Fen Fak. Mecm., (B)20(1-2): 49-94.
    • O'Reilly, M.G. 1995. A new genus of copepod (Copepoda: Poecilostomatoida) commensal with the maldanid polychaete Rhodine gracilior, with a review of the family Clausiidae. J. nat. Hist. 29: 47-64.
    • Por, F.D., 1964. A study of Levantine and Pontic Harpacticoida (Crustacea, Copepoda). - Zool. Verh., Leiden, 64: 1-128.
    • Raibaut, A., Combes, C. & Benoit, F. 1998. Analysis of the parasitic copepod species richness among Mediterranean fish. J. mar. Syst. 15: 185-206.
    • Razouls, C. 1995. Diversité et répartition géographique chez les copépodes pélagiques. 1. Calanoida. Annal. Inst. Oceanogr. 71: 81-404.
    • Razouls, C. 1995. Diversité et répartition géographique chez les copépodes pélagiques. 2. Platycopioida, Misophrioida, Mormonilloida, Cyclopoida, Poecilostomatoida, Siphonostomatoida, Harpacticoida, Monstrilloida. Annal. Inst. Oceanogr. 72: 1-149.
    • Soyer, J., 1971. Bionomie benthique du plateau continental de la côte catalane française. III. Les peuplements de Copépodes Harpacticoïdes (Crustacea). - Vie Milieu, (B)21(2): 337-511.
    • Zagami, G., Campolmi, M, & Costanzo, G. 2000. A new species of Stephos T.Scott, 1892 (Copepoda: Calanoida) from coastal waters of Sicily, Italy. J. plankton Res. 22: 15-27.

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